The tribe that claims this prize can reshape the world any way they see fit. The Amanitas Empire and The Lepiota Order see the myth as a chance to not only defeat the other nations (the Bolete Tribe and the Morel Brotherhood) but also wipe out all of their enemies and put their tribes at the forefront of terrestrial evolution.

Even while the Spore War rages, a great quest amongst all four tribes begins: a race to uncover the truth and reshape the evolutionary course of the Earth. As for the humans, they continue to bumble their way through the world completely unaware of the diminutive forces rising against them. Pax, a Bolete mushroom from the outskirts, doesn't yet understand his place in the events to come. He'll have to learn, and fast, that he’s unwittingly been thrust into the middle of this situation.